I am new to the forum but have been into wildlife photography, mostly birds, for a while. Originally, I was a Nikon DSLR user, then a Sony Mirrorless user, and since the Z8, I started migrating to Nikon Z for most of my photography needs.
For birding in my area, reach usually becomes the issue, and I have stuck with my Sony 200-600mm + TC 1.4 on my Sony a7rIV. I was strongly considering getting the Nikon 800mm f6.3 PF, but one thing that held me back was the 30% increase in resolution the a7rIV has over the Z8.
Now, with Sony anoucing the 400-800mm, I am more confused about which way to go. The way I see it:
The 200-600mm f6.3 gets me:
200-600mm f6.3
+ TC 1.4x 280-840mm f9
+ TC 2x 400-1,200 f13
The 400-800mm f8 gets me:
400-800 f8
+ TC 1.4x 560-1,120 f11
+ TC 2x 800-1,600mm f16
The 800mm f6.3 PF gets me:
800mm f6.3
+ TC 1.4x 1,120mm f9
+ TC 2x 1,600mm f13
Based on my experience with the TC 2x on the 200-600mm at f13, I am not sure the TC 2x on the 400-800mm at f16 will be useful for much outside of video. So I feel that for max optical reach, the 800mm PF wins. Also, in lower light, the 800mm PF wins being 2/3 of a stop faster, but the Sony 400-800mm has the ability to zoom, which is very convenient. The Sony will also have the camera resolution advantage for cropping. I wish Sony allowed Sigma to make TCs for the 300-600mm f4 because that sounds like the perfect wildlife lens for me if it supported TCs.
I know the 400-800mm is not out yet, but based on what we know so far, which lens would you go for the Sony 400-800mm f8 or the Nikon 800mm PF f6.3?
I'm a Nikon shooter but have to admit that new Sony 400-800 is a very interesting lens.
To address your concern about the difference in resultion between the A7R IV and Z8, yes the A7 IV has 35% more pixels. However, that translates to a 16% increase in resultion. (The square root of the ratio is what determines the difference) So, while there is a difference, it's one that would only be noticeable when comparing photos at 100%.
Since the lens you currently use and one of the two you're considering are zooms, I'd ask if you'll miss the shorter focal lengths. One of the reasons I upgraded from a Nikkor 200-500 to the 800 PF is that a review of the focal lengths used to make my favorite photos with the zoom revealed that 90% were made at 500mm.
I used that lens with an APS-C D500 so, the 500mm lens translates to a full frame equivalent 750mm angle of view. If your photography with the 200-600 is similarly biased toward the 600mm end, moving to a prime could be a good fit. If, say, 1/3 or more of your favorites with that lens have been made at 500mm or shorter, you might find a prime lens not such a good fit.
By the same token, if a significant percentage of your favs have been made in the 200-350mm range, you might find the 400-800 too long. Keeping a shorter prime or zoom in your arsenal to complement the 400-800 could be a good strategy to address that need.
What intrigues me about the 400-800 is the 100mm diameter entrance pupil at 800mm f/8. It's among the few lenses with an entrance pupil that large. Most lenses with larger apertures cost 2x to 4x as much.
At 800mm f8, it has an entrance pupil just a skosh larger (100mm vs 95mm) than the 200-600 at 600mm f/6.3. Adding a 1.4x TC to the 200-600, you achieve the same angle of view. With both lenses having very similar entrance pupils, they'll make photos having similar noise levels and depths of field.
Personally, I'd suggest looking for reviews comparing the two for sharpness and detail rendering. If the 400-800 is a touch sharper at its native 800mm than the 200-600 when cropped or paired with a TC to match the framing, that could translate to the longer zoom having an advantage.
I resist recommending system changes. They can be expensive and potentially frustrating if one doesn't like the new user interface. That said and since you're considering the 800 PF, that lens does collect about 60% more light from a subject in the frame than the 200-600 or 400-800. That should translate to a noticeable reduction in image noise at the same shutter speed. It's also likely to be at least as sharp and probably a skosh sharper. Plus, it's not too heavy; about the same weight as the 400-800.
Good luck with your decision.