Not to invite an argument but I think you might be a bit alarmist. Personally, I don't see any significant "gaps" in their product line for wildlife and nature photography, but what the heck do I know? I'm just an amateur
What I do see are the major camera manufacturers regularly leap-frogging one another with innovation, with the lead frog periodically flagging when their rivals turn up the temperature. A couple of years ago I read and heard how a lot of wildlife photographers were abandoning Nikon for Sony, and now Nikon is one of the (if not THE) industry leaders. Lots of unnecessary teeth gnashing and hand wringing, IMHO.